ERDDAP > info > bcodmo_dataset_750328

Grid DAP Data	Sub- set	Table DAP Data	Make A Graph	W M S	Source Data Files	Acces- sible	Title	Sum- mary	ISO, Metadata	Back- ground Info	RSS	E mail	Institution	Dataset ID
		data	graph		files	public	Quantification of a previously undescribed fast-start of larval clownfish targeting copepod prey: predatory posture, speed and acceleration from high-speed video, July 2015		I   M	background			BCO-DMO	bcodmo_dataset_750328

The Dataset's Variables and Attributes

Row Type	Variable Name	Attribute Name	Data Type	Value
attribute	NC_GLOBAL	access_formats	String	.htmlTable,.csv,.json,.mat,.nc,.tsv
attribute	NC_GLOBAL	acquisition_description	String	Permits:\u00a0All protocols and experiments, described below, followed institutional guidelines and were approved by the University of Hawaii Institutional Animal Care & Use Committee (IACUC protocol number 2099). Experimental design:\u00a0We set three larval fish age-classes of the clownfish, Amphiprion ocellaris (early: 1 to 5 days post-hatch [dph]; mid: 6 to 9 dph; and late: 11 to 14 dph) against three developmental stages of the copepod prey, Bestiolina similis (nauplii: NIII-NIV stages; copepodites: CII- CIII stages; and adults: CVI stage). The choice of developmental stages provided a range in prey size (length: ~100 to 500 \u00b5m; McKinnon et al., 2003), mechanosensitivity, and escape performance (Buskey et al., 2017). We designed the experiment to quantify how strike posture changed through larval development (and size: ~4 to 8 mm total length; Jackson and Lenz, 2016), while also assessing the effect of a prey\u2019s stage on its predator\u2019s posture. Larval fish-rearing conditions: The experiments used larval clownfish lab- reared over their two-week planktonic phase. Their rearing, the culturing of copepod prey, the experimental apparatus and protocols, and the high-speed video recording and analysis software have been described previously (Robinson et al., accepted). Briefly, up to 200 recently hatched larvae were raised in a 30-L seawater aquarium kept at 24-26\u00b0 C on a 12:12 L:D light cycle. They were fed twice daily on a mixed diet of rotifers (Brachionus plicatilis) and different developmental stages of another calanoid copepod (Parvocalanus crassirostris). Different prey were used for daily feeding than for experiments so that fish were exposed to a novel prey type during their trial, thus avoiding complications arising from learned feeding behavior and laboratory acclimation. Behavioral observations and video set-up:\u00a0For the experiments, two larvae that had been kept without food for 4 to 6 hours were placed into a circular observation chamber of 20 cm diameter, filled with seawater to a depth of 2 cm containing copepods at a density of 0.2 to 0.7 individuals ml-1. Experimental trials lasted for one hour or less and no fish larvae were used in more than one trial. Interactions between fish larvae and copepods were recorded at 500 frames per second (fps) using a Photron FastCAM SA4 video camera mounted above the observation chamber with dark-field illumination. The field-of-view of the camera was 35 x 35 mm with an image resolution of 1024 x 1024 pixels. Data analysis:\u00a0Predatory attacks were analyzed to characterize the temporal sequence up to and through the strike of fish larvae on different copepod developmental stages. We quantified the duration (in sec) of the approach phase of the fish, defined and described in Robinson et al. (accepted) as beginning when the tail stopped beating and started to bend to the left or right, and ending at t0. Thirty-seven successful captures (n = 37) by fish larvae were analyzed using the Fiji software package built on ImageJ (v1.51) (Schindelin et al., 2012) to digitize the changing body shape of the fish as it prepared for and initiated the strike. For each interaction, we established a temporal reference (t0) as the image just prior to the opening of the fish\u2019s mouth. Twenty-five frames, including 12 frames before and 12 frames after t0, were then extracted and the posture of the fish larva was characterized frame-by-frame. Twelve frames (24 ms) before t0, labeled as t-24, was chosen as a standardized interval that captured the final approach phase of fish in our trials, which ranged from 28 to 1130 ms preceding t0. The median speed during the final frame of approach (t-2) was 0 mm s-1. Therefore, fish were most often motionless at this time interval, meaning that all motion involved in the acceleration of the strike occurred after t0. Twelve frames after t0, labelled as t+24, was chosen as a standardized interval as it always included peak strike speed, copepod capture, and subsequent deceleration of the fish. Starting with t-24 and continuing every other frame up to and including t+24, we digitized the x,y coordinates of 12 points along the central axis of the larva. Because its position with respect to the fish did not change, we used a small pigment spot located at the narrowest point between the eyes as a spatial reference for each frame (origin at x=0, y=0). The position of the copepod in each frame was also digitized to obtain the change in distance over time between fish and copepod.\u00a0 We quantified the relative curvature of the body and caudal fin during the larva\u2019s initial lunge, frame-by-frame from t0 to t+8 (8 ms, i.e., 4 frames after t0). To do so, we used the oval tool in Fiji to place a circle within the curl of the tail. The \u201cfit circle\u201d and \u201cmeasure\u201d commands were then used to calculate the area (A) of the circle. From the circle\u2019s area, we derived the reciprocal radius, r -1 = \u221a(\u03c0 / A) as a metric of relative curvature of the larva\u2019s caudal fin, with greater values being more curved and lesser values being less curved. The measurement of reciprocal radius became less reliable as an estimate of curvature after t+8 because the tails of most fish began to curve in the opposing direction, making the inscribed circle too large to measure. We therefore employed another relative measure of curvature, the straight-line distance between the tip of the tail and the narrowest point between the eyes (chord length), divided by the length of the fish when its body was straight (fish length), also measured between the tip of the tail and origin/pigment. These normalized distances (chord length-to-fish length ratio, or CVF) approached 1 when the fish was straight and were decreasing fractions of 1 when the fish was increasingly bent into a J-like posture. To measure speed of the fish during its predatory lunge (in mm s-1), we tracked the changing position of the pigment between its eyes from t-4 to t+20 (4 ms, i.e., 2 frames before t0 and 20 ms, i.e., 10 frames after t0, respectively) and divided the frame-by-frame distance travelled by the time elapsed between frames. In addition, the distance from the spot between the eyes and the tip of the mouth was measured at t0 and t+8 to determine the contribution of jaw extension to prey capture.\u00a0
attribute	NC_GLOBAL	awards_0_award_nid	String	562096
attribute	NC_GLOBAL	awards_0_award_number	String	OCE-1235549
attribute	NC_GLOBAL	awards_0_data_url	String	http://www.nsf.gov/awardsearch/showAward.do?AwardNumber=1235549
attribute	NC_GLOBAL	awards_0_funder_name	String	NSF Division of Ocean Sciences
attribute	NC_GLOBAL	awards_0_funding_acronym	String	NSF OCE
attribute	NC_GLOBAL	awards_0_funding_source_nid	String	355
attribute	NC_GLOBAL	awards_0_program_manager	String	David L. Garrison
attribute	NC_GLOBAL	awards_0_program_manager_nid	String	50534
attribute	NC_GLOBAL	cdm_data_type	String	Other
attribute	NC_GLOBAL	comment	String	Clownfish predatory posture Quantification of a previously undescribed fast-start of larval clownfish targeting copepod prey PI's: P. Lenz, D. Hartline (U Hawaii - Manoa) version: 2018-11-28
attribute	NC_GLOBAL	Conventions	String	COARDS, CF-1.6, ACDD-1.3
attribute	NC_GLOBAL	creator_email	String	info at bco-dmo.org
attribute	NC_GLOBAL	creator_name	String	BCO-DMO
attribute	NC_GLOBAL	creator_type	String	institution
attribute	NC_GLOBAL	creator_url	String	https://www.bco-dmo.org/
attribute	NC_GLOBAL	data_source	String	extract_data_as_tsv version 2.3 19 Dec 2019
attribute	NC_GLOBAL	date_created	String	2018-11-28T19:50:41Z
attribute	NC_GLOBAL	date_modified	String	2019-03-18T14:51:15Z
attribute	NC_GLOBAL	defaultDataQuery	String	&time<now
attribute	NC_GLOBAL	doi	String	10.1575/1912/bco-dmo.750328.1
attribute	NC_GLOBAL	infoUrl	String	https://www.bco-dmo.org/dataset/750328
attribute	NC_GLOBAL	institution	String	BCO-DMO
attribute	NC_GLOBAL	instruments_0_acronym	String	camera
attribute	NC_GLOBAL	instruments_0_dataset_instrument_description	String	A Photron FastCAM SA4 high-speed video camera with a Nikon micro-NIKKOR 60 mm lens and 36 mm extension tube was used to record predator-prey interaction. The experimental chamber was illuminated by a dark field, ring light, Fiber-Lite MI-150 high-intensity illuminator, Dolan-Jenner. The camera was mounted on a manually-operated, linear positioning slide (Automation Gages Inc.)
attribute	NC_GLOBAL	instruments_0_dataset_instrument_nid	String	750335
attribute	NC_GLOBAL	instruments_0_description	String	All types of photographic equipment including stills, video, film and digital systems.
attribute	NC_GLOBAL	instruments_0_instrument_external_identifier	String	https://vocab.nerc.ac.uk/collection/L05/current/311/
attribute	NC_GLOBAL	instruments_0_instrument_name	String	Camera
attribute	NC_GLOBAL	instruments_0_instrument_nid	String	520
attribute	NC_GLOBAL	instruments_0_supplied_name	String	high-speed video camera
attribute	NC_GLOBAL	keywords	String	2ms, accelation, acceleration, ACCELERATION_T0, ACCELERATION_T10ms, ACCELERATION_T16ms, ACCELERATION_T1point5ms, ACCELERATION_T20ms, ACCELERATION_T2ms, ACCELERATION_T4ms, ACCELERATION_T6ms, ACCELERATION_T8ms, age, approach, APPROACH_DURATION, array, array-data, bco, bco-dmo, biological, birthdate, body, BODY_LENGTH, capture, category, chemical, class, clip, CLIP_DURATION, CLIP_END, CLIP_ID, CLIP_START, comprehensive, cvf, CVF_T0, CVF_T10ms, CVF_T12ms, CVF_T14ms, CVF_T16ms, CVF_T18ms, CVF_T20ms, CVF_T2ms, CVF_T4ms, CVF_T6ms, CVF_T8ms, data, dataset, date, distance, dmo, dph, duration, end, erddap, fish, FISH_AGE_CLASS, frame, FRAME_APPROACH_START, FRAME_T0_STRIKE, inverse, INVERSE_RADIUS_T0, INVERSE_RADIUS_T2ms, INVERSE_RADIUS_T4ms, INVERSE_RADIUS_T6ms, INVERSE_RADIUS_T8ms, large, length, management, oceanography, office, peak, PEAK_ACCELERATION, PEAK_STRIKE_SPEED, pixel, PIXEL_TO_MM, preliminary, prey, PREY_STAGE_CLASS, radius, speed, stage, STAGE_CATEGORY, start, stewardship, strike, STRIKE_DISTANCE_MM, STRIKE_DISTANCE_PIXEL, STRIKE_SPEED_T11ms, STRIKE_SPEED_T14ms, STRIKE_SPEED_T18ms, STRIKE_SPEED_T1ms, STRIKE_SPEED_T22ms, STRIKE_SPEED_T3ms, STRIKE_SPEED_T5ms, STRIKE_SPEED_T7ms, STRIKE_SPEED_T9ms, STRIKE_SPEED_T_2ms, system, t10ms, t11ms, t12ms, t14ms, t16ms, t18ms, t1ms, t1point5ms, t20ms, t22ms, t2ms, t3ms, t4ms, t5ms, t6ms, t7ms, t8ms, t9ms, time, TIME_FROM_T0_TO_CAPTURE, TIME_TO_PEAK_ACCELATION, trial
attribute	NC_GLOBAL	license	String	https://www.bco-dmo.org/dataset/750328/license
attribute	NC_GLOBAL	metadata_source	String	https://www.bco-dmo.org/api/dataset/750328
attribute	NC_GLOBAL	param_mapping	String	{'750328': {}}
attribute	NC_GLOBAL	parameter_source	String	https://www.bco-dmo.org/mapserver/dataset/750328/parameters
attribute	NC_GLOBAL	people_0_affiliation	String	University of Hawaii at Manoa
attribute	NC_GLOBAL	people_0_affiliation_acronym	String	PBRC
attribute	NC_GLOBAL	people_0_person_name	String	Dr Petra Lenz
attribute	NC_GLOBAL	people_0_person_nid	String	473047
attribute	NC_GLOBAL	people_0_role	String	Principal Investigator
attribute	NC_GLOBAL	people_0_role_type	String	originator
attribute	NC_GLOBAL	people_1_affiliation	String	University of Hawaii at Manoa
attribute	NC_GLOBAL	people_1_affiliation_acronym	String	PBRC
attribute	NC_GLOBAL	people_1_person_name	String	Dr Daniel K Hartline
attribute	NC_GLOBAL	people_1_person_nid	String	562100
attribute	NC_GLOBAL	people_1_role	String	Co-Principal Investigator
attribute	NC_GLOBAL	people_1_role_type	String	originator
attribute	NC_GLOBAL	people_2_affiliation	String	Woods Hole Oceanographic Institution
attribute	NC_GLOBAL	people_2_affiliation_acronym	String	WHOI BCO-DMO
attribute	NC_GLOBAL	people_2_person_name	String	Nancy Copley
attribute	NC_GLOBAL	people_2_person_nid	String	50396
attribute	NC_GLOBAL	people_2_role	String	BCO-DMO Data Manager
attribute	NC_GLOBAL	people_2_role_type	String	related
attribute	NC_GLOBAL	project	String	PreyEscape
attribute	NC_GLOBAL	projects_0_acronym	String	PreyEscape
attribute	NC_GLOBAL	projects_0_description	String	Description from NSF award abstract: This study will experimentally elucidate the dynamics of predator evasion by different species and life stages of copepod responding to a model larval fish predator. The PIs will use standard and high-speed videographic and cutting-edge holographic techniques. Predator-prey interactions within planktonic communities are key to understanding how energy is transferred within complex marine food webs. Of particular interest are those between the highly numerous copepods and one of their more important predators, the ichthyoplankton (the planktonic larval stages of fishes). The larvae of most fishes are planktivorous and heavily dependent on copepods for food. In general, evasion success increases with age in copepods and decreases with the age of the fish predator. How this plays out in detail is critical in determining predatory attack outcomes and the effect these have on predator and prey survival. To address this problem, different copepod developmental stages will be tested against several levels of predator competence, and the results examined for: 1) the success or failure of attacks for different combinations of predator and prey age class; 2) the kinematics (reaction latencies and trajectory orientation) for escape attempts, successful and unsuccessful, for different age classes of copepod; 3) the hydrodynamic cues generated by different ages and attack strategies of the predator and the sensitivity of different prey stages to these cues; and 4) the success or failure of the predatory approach and attack strategies at each prey stage. The data obtained will be used to inform key issues of zooplankton population dynamics. For the prey these include: predator-evasion capabilities and importance of detection ability, reaction speed, escape speed, escape orientation, and trajectory irregularity; for the predator they are: capabilities and importance of mouth gape size, stealthiness, hydrodynamic disturbance production, and lunge kinematics.
attribute	NC_GLOBAL	projects_0_end_date	String	2016-08
attribute	NC_GLOBAL	projects_0_geolocation	String	Pacific
attribute	NC_GLOBAL	projects_0_name	String	The Drive to Survive: Copepods vs Ichthyoplankton
attribute	NC_GLOBAL	projects_0_project_nid	String	562097
attribute	NC_GLOBAL	projects_0_start_date	String	2012-09
attribute	NC_GLOBAL	publisher_name	String	Biological and Chemical Oceanographic Data Management Office (BCO-DMO)
attribute	NC_GLOBAL	publisher_type	String	institution
attribute	NC_GLOBAL	sourceUrl	String	(local files)
attribute	NC_GLOBAL	standard_name_vocabulary	String	CF Standard Name Table v55
attribute	NC_GLOBAL	summary	String	Posture, strike speed, and acceleration of clownfish larval attack on copepods, from high-speed videos, June-July 2015. Results of these data are published in Fashingbauer et al (in revision, J. Exp. Biol.) and Robinson et al (accepted, MEPS).
attribute	NC_GLOBAL	title	String	Quantification of a previously undescribed fast-start of larval clownfish targeting copepod prey: predatory posture, speed and acceleration from high-speed video, July 2015
attribute	NC_GLOBAL	version	String	1
attribute	NC_GLOBAL	xml_source	String	osprey2erddap.update_xml() v1.3
variable	TRIAL_DATE		String
attribute	TRIAL_DATE	bcodmo_name	String	date
attribute	TRIAL_DATE	description	String	date the experimental observations took place formatted as yyyy-mm-dd
attribute	TRIAL_DATE	long_name	String	TRIAL DATE
attribute	TRIAL_DATE	nerc_identifier	String	https://vocab.nerc.ac.uk/collection/P01/current/ADATAA01/
attribute	TRIAL_DATE	source_name	String	TRIAL_DATE
attribute	TRIAL_DATE	time_precision	String	1970-01-01
attribute	TRIAL_DATE	units	String	unitless
variable	BIRTHDATE		String
attribute	BIRTHDATE	bcodmo_name	String	date
attribute	BIRTHDATE	description	String	birthdate of fish; when the fish hatched formatted as yyyy-mm-dd
attribute	BIRTHDATE	long_name	String	BIRTHDATE
attribute	BIRTHDATE	nerc_identifier	String	https://vocab.nerc.ac.uk/collection/P01/current/ADATAA01/
attribute	BIRTHDATE	time_precision	String	1970-01-01
attribute	BIRTHDATE	units	String	unitless
variable	DPH		byte
attribute	DPH	_FillValue	byte	127
attribute	DPH	actual_range	byte	1, 13
attribute	DPH	bcodmo_name	String	age
attribute	DPH	description	String	larval fish age; days post hatch
attribute	DPH	long_name	String	DPH
attribute	DPH	units	String	days post hatch
variable	FISH_AGE_CLASS		String
attribute	FISH_AGE_CLASS	bcodmo_name	String	age
attribute	FISH_AGE_CLASS	description	String	larval age-group of Amphiprion ocellaris: early (1-5 dph [days post-hatch]); mid (6-9 dph); or late (11-14 dph)
attribute	FISH_AGE_CLASS	long_name	String	FISH AGE CLASS
attribute	FISH_AGE_CLASS	units	String	unitless
variable	PREY_STAGE_CLASS		String
attribute	PREY_STAGE_CLASS	bcodmo_name	String	stage
attribute	PREY_STAGE_CLASS	description	String	developmental stage-class of Bestiolina similis: nauplii (NIII-NIV stages); early copepodites (CII-CIII stages; called just "copepodites" in hard-drive folders); late copepodites (CV stage); adults (CVI stage)
attribute	PREY_STAGE_CLASS	long_name	String	PREY STAGE CLASS
attribute	PREY_STAGE_CLASS	units	String	unitless
variable	STAGE_CATEGORY		String
attribute	STAGE_CATEGORY	bcodmo_name	String	stage
attribute	STAGE_CATEGORY	description	String	predator-prey age group combination
attribute	STAGE_CATEGORY	long_name	String	STAGE CATEGORY
attribute	STAGE_CATEGORY	units	String	unitless
variable	CLIP_ID		String
attribute	CLIP_ID	bcodmo_name	String	sample
attribute	CLIP_ID	description	String	two-letter identifier in TIF file name - separating clips
attribute	CLIP_ID	long_name	String	CLIP ID
attribute	CLIP_ID	nerc_identifier	String	https://vocab.nerc.ac.uk/collection/P02/current/ACYC/
attribute	CLIP_ID	units	String	unitless
variable	CLIP_START		short
attribute	CLIP_START	_FillValue	short	32767
attribute	CLIP_START	actual_range	short	1, 31824
attribute	CLIP_START	bcodmo_name	String	unknown
attribute	CLIP_START	description	String	first frame in clip; numbered with reference to other clips in the same data folder
attribute	CLIP_START	long_name	String	CLIP START
attribute	CLIP_START	units	String	unitless
variable	CLIP_END		int
attribute	CLIP_END	_FillValue	int	2147483647
attribute	CLIP_END	actual_range	int	539, 40554
attribute	CLIP_END	bcodmo_name	String	unknown
attribute	CLIP_END	description	String	final frame in clip; numbered with reference to other clips in the same data folder
attribute	CLIP_END	long_name	String	CLIP END
attribute	CLIP_END	units	String	unitless
variable	CLIP_DURATION		short
attribute	CLIP_DURATION	_FillValue	short	32767
attribute	CLIP_DURATION	actual_range	short	539, 9561
attribute	CLIP_DURATION	bcodmo_name	String	duration
attribute	CLIP_DURATION	description	String	length of a clip
attribute	CLIP_DURATION	long_name	String	CLIP DURATION
attribute	CLIP_DURATION	units	String	# of frames
variable	PIXEL_TO_MM		float
attribute	PIXEL_TO_MM	_FillValue	float	NaN
attribute	PIXEL_TO_MM	actual_range	float	28.716, 29.186
attribute	PIXEL_TO_MM	bcodmo_name	String	unknown
attribute	PIXEL_TO_MM	description	String	calibration ratio of the number of pixels per millimeter; unique for each trial date
attribute	PIXEL_TO_MM	long_name	String	PIXEL TO MM
attribute	PIXEL_TO_MM	units	String	pixels per millimeter
variable	BODY_LENGTH		float
attribute	BODY_LENGTH	_FillValue	float	NaN
attribute	BODY_LENGTH	actual_range	float	4.8, 10.3
attribute	BODY_LENGTH	bcodmo_name	String	length
attribute	BODY_LENGTH	description	String	body length; measured as total length
attribute	BODY_LENGTH	long_name	String	BODY LENGTH
attribute	BODY_LENGTH	units	String	milllimeters
variable	FRAME_APPROACH_START		short
attribute	FRAME_APPROACH_START	_FillValue	short	32767
attribute	FRAME_APPROACH_START	actual_range	short	183, 5700
attribute	FRAME_APPROACH_START	bcodmo_name	String	unknown
attribute	FRAME_APPROACH_START	description	String	frame number when final approach began (per definition set by Robinson et al. (accepted))
attribute	FRAME_APPROACH_START	long_name	String	FRAME APPROACH START
attribute	FRAME_APPROACH_START	units	String	frame #
variable	APPROACH_DURATION		float
attribute	APPROACH_DURATION	_FillValue	float	NaN
attribute	APPROACH_DURATION	actual_range	float	0.028, 1.13
attribute	APPROACH_DURATION	bcodmo_name	String	duration
attribute	APPROACH_DURATION	description	String	duration of the final approach (per definition set by Robinson et al. (accepted))
attribute	APPROACH_DURATION	long_name	String	APPROACH DURATION
attribute	APPROACH_DURATION	units	String	seconds
variable	FRAME_T0_STRIKE		short
attribute	FRAME_T0_STRIKE	_FillValue	short	32767
attribute	FRAME_T0_STRIKE	actual_range	short	325, 5841
attribute	FRAME_T0_STRIKE	bcodmo_name	String	unknown
attribute	FRAME_T0_STRIKE	description	String	frame number at t0; moment when mouth began to open
attribute	FRAME_T0_STRIKE	long_name	String	FRAME T0 STRIKE
attribute	FRAME_T0_STRIKE	units	String	frame #
variable	STRIKE_DISTANCE_PIXEL		float
attribute	STRIKE_DISTANCE_PIXEL	_FillValue	float	NaN
attribute	STRIKE_DISTANCE_PIXEL	actual_range	float	9.564, 59.796
attribute	STRIKE_DISTANCE_PIXEL	bcodmo_name	String	unknown
attribute	STRIKE_DISTANCE_PIXEL	description	String	distance at t0 between edge of fish mouth and rostrum of copepod (or center of copepod if rostrum not visible)
attribute	STRIKE_DISTANCE_PIXEL	long_name	String	STRIKE DISTANCE PIXEL
attribute	STRIKE_DISTANCE_PIXEL	units	String	pixels
variable	STRIKE_DISTANCE_MM		float
attribute	STRIKE_DISTANCE_MM	_FillValue	float	NaN
attribute	STRIKE_DISTANCE_MM	actual_range	float	0.332, 2.069
attribute	STRIKE_DISTANCE_MM	bcodmo_name	String	unknown
attribute	STRIKE_DISTANCE_MM	description	String	distance at t0 between edge of fish mouth and rostrum of copepod (or center of copepod if rostrum not visible)
attribute	STRIKE_DISTANCE_MM	long_name	String	STRIKE DISTANCE MM
attribute	STRIKE_DISTANCE_MM	units	String	milllimeters
variable	TIME_FROM_T0_TO_CAPTURE		byte
attribute	TIME_FROM_T0_TO_CAPTURE	_FillValue	byte	127
attribute	TIME_FROM_T0_TO_CAPTURE	actual_range	byte	4, 14
attribute	TIME_FROM_T0_TO_CAPTURE	bcodmo_name	String	time_elapsed
attribute	TIME_FROM_T0_TO_CAPTURE	description	String	time from t0 until capture (copepod completely in fish mouth)
attribute	TIME_FROM_T0_TO_CAPTURE	long_name	String	TIME FROM T0 TO CAPTURE
attribute	TIME_FROM_T0_TO_CAPTURE	nerc_identifier	String	https://vocab.nerc.ac.uk/collection/P01/current/ELTMZZZZ/
attribute	TIME_FROM_T0_TO_CAPTURE	units	String	milliseconds
variable	PEAK_STRIKE_SPEED		float
attribute	PEAK_STRIKE_SPEED	_FillValue	float	NaN
attribute	PEAK_STRIKE_SPEED	actual_range	float	88.785, 394.522
attribute	PEAK_STRIKE_SPEED	bcodmo_name	String	unknown
attribute	PEAK_STRIKE_SPEED	description	String	maximum speed attained by clownfish from t-2ms to t22ms
attribute	PEAK_STRIKE_SPEED	long_name	String	PEAK STRIKE SPEED
attribute	PEAK_STRIKE_SPEED	units	String	millimeters/second
variable	STRIKE_SPEED_T_2ms		float
attribute	STRIKE_SPEED_T_2ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T_2ms	actual_range	float	0.0, 19.393
attribute	STRIKE_SPEED_T_2ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T_2ms	description	String	speed of fish during attack at time = -2 milliseconds
attribute	STRIKE_SPEED_T_2ms	long_name	String	STRIKE SPEED T 2ms
attribute	STRIKE_SPEED_T_2ms	units	String	millimeters/second
variable	STRIKE_SPEED_T1ms		float
attribute	STRIKE_SPEED_T1ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T1ms	actual_range	float	0.0, 62.486
attribute	STRIKE_SPEED_T1ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T1ms	description	String	speed of fish during attack at time = 1 millisecond
attribute	STRIKE_SPEED_T1ms	long_name	String	STRIKE SPEED T1ms
attribute	STRIKE_SPEED_T1ms	units	String	millimeters/second
variable	STRIKE_SPEED_T3ms		float
attribute	STRIKE_SPEED_T3ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T3ms	actual_range	float	0.0, 247.711
attribute	STRIKE_SPEED_T3ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T3ms	description	String	speed of fish during attack at time = 3 milliseconds
attribute	STRIKE_SPEED_T3ms	long_name	String	STRIKE SPEED T3ms
attribute	STRIKE_SPEED_T3ms	units	String	millimeters/second
variable	STRIKE_SPEED_T5ms		float
attribute	STRIKE_SPEED_T5ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T5ms	actual_range	float	17.15, 318.619
attribute	STRIKE_SPEED_T5ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T5ms	description	String	speed of fish during attack at time = 5 milliseconds
attribute	STRIKE_SPEED_T5ms	long_name	String	STRIKE SPEED T5ms
attribute	STRIKE_SPEED_T5ms	units	String	millimeters/second
variable	STRIKE_SPEED_T7ms		float
attribute	STRIKE_SPEED_T7ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T7ms	actual_range	float	71.792, 394.522
attribute	STRIKE_SPEED_T7ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T7ms	description	String	speed of fish during attack at time = 7 milliseconds
attribute	STRIKE_SPEED_T7ms	long_name	String	STRIKE SPEED T7ms
attribute	STRIKE_SPEED_T7ms	units	String	millimeters/second
variable	STRIKE_SPEED_T9ms		float
attribute	STRIKE_SPEED_T9ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T9ms	actual_range	float	54.485, 336.499
attribute	STRIKE_SPEED_T9ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T9ms	description	String	speed of fish during attack at time = 9 milliseconds
attribute	STRIKE_SPEED_T9ms	long_name	String	STRIKE SPEED T9ms
attribute	STRIKE_SPEED_T9ms	units	String	millimeters/second
variable	STRIKE_SPEED_T11ms		float
attribute	STRIKE_SPEED_T11ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T11ms	actual_range	float	24.423, 210.475
attribute	STRIKE_SPEED_T11ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T11ms	description	String	speed of fish during attack at time = 11 milliseconds
attribute	STRIKE_SPEED_T11ms	long_name	String	STRIKE SPEED T11ms
attribute	STRIKE_SPEED_T11ms	units	String	millimeters/second
variable	STRIKE_SPEED_T14ms		float
attribute	STRIKE_SPEED_T14ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T14ms	actual_range	float	35.52, 155.948
attribute	STRIKE_SPEED_T14ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T14ms	description	String	speed of fish during attack at time = 14 milliseconds
attribute	STRIKE_SPEED_T14ms	long_name	String	STRIKE SPEED T14ms
attribute	STRIKE_SPEED_T14ms	units	String	millimeters/second
variable	STRIKE_SPEED_T18ms		float
attribute	STRIKE_SPEED_T18ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T18ms	actual_range	float	31.19, 149.085
attribute	STRIKE_SPEED_T18ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T18ms	description	String	speed of fish during attack at time = 18 milliseconds
attribute	STRIKE_SPEED_T18ms	long_name	String	STRIKE SPEED T18ms
attribute	STRIKE_SPEED_T18ms	units	String	millimeters/second
variable	STRIKE_SPEED_T22ms		float
attribute	STRIKE_SPEED_T22ms	_FillValue	float	NaN
attribute	STRIKE_SPEED_T22ms	actual_range	float	0.0, 120.694
attribute	STRIKE_SPEED_T22ms	bcodmo_name	String	unknown
attribute	STRIKE_SPEED_T22ms	description	String	speed of fish during attack at time = 22 milliseconds
attribute	STRIKE_SPEED_T22ms	long_name	String	STRIKE SPEED T22ms
attribute	STRIKE_SPEED_T22ms	units	String	millimeters/second
variable	TIME_TO_PEAK_ACCELATION		byte
attribute	TIME_TO_PEAK_ACCELATION	_FillValue	byte	127
attribute	TIME_TO_PEAK_ACCELATION	actual_range	byte	2, 8
attribute	TIME_TO_PEAK_ACCELATION	bcodmo_name	String	time_elapsed
attribute	TIME_TO_PEAK_ACCELATION	description	String	time from t0 until peak acceleration is reached
attribute	TIME_TO_PEAK_ACCELATION	long_name	String	TIME TO PEAK ACCELATION
attribute	TIME_TO_PEAK_ACCELATION	nerc_identifier	String	https://vocab.nerc.ac.uk/collection/P01/current/ELTMZZZZ/
attribute	TIME_TO_PEAK_ACCELATION	units	String	milliseconds
variable	PEAK_ACCELERATION		double
attribute	PEAK_ACCELERATION	_FillValue	double	NaN
attribute	PEAK_ACCELERATION	actual_range	double	18274.145, 115205.267
attribute	PEAK_ACCELERATION	bcodmo_name	String	unknown
attribute	PEAK_ACCELERATION	description	String	maximum acceleration from t0 to t20ms
attribute	PEAK_ACCELERATION	long_name	String	PEAK ACCELERATION
attribute	PEAK_ACCELERATION	units	String	millimeters/second^2
variable	ACCELERATION_T0		double
attribute	ACCELERATION_T0	_FillValue	double	NaN
attribute	ACCELERATION_T0	actual_range	double	-5756.496, 20828.791
attribute	ACCELERATION_T0	bcodmo_name	String	unknown
attribute	ACCELERATION_T0	description	String	acceleration of fish at t0
attribute	ACCELERATION_T0	long_name	String	ACCELERATION T0
attribute	ACCELERATION_T0	units	String	millimeters/second^2
variable	ACCELERATION_T2ms		double
attribute	ACCELERATION_T2ms	_FillValue	double	NaN
attribute	ACCELERATION_T2ms	actual_range	double	-17330.602, 115205.267
attribute	ACCELERATION_T2ms	bcodmo_name	String	unknown
attribute	ACCELERATION_T2ms	description	String	acceleration of fish at t2ms
attribute	ACCELERATION_T2ms	long_name	String	ACCELERATION T2ms
attribute	ACCELERATION_T2ms	units	String	millimeters/second^2
variable	ACCELERATION_T4ms		double
attribute	ACCELERATION_T4ms	_FillValue	double	NaN
attribute	ACCELERATION_T4ms	actual_range	double	-9242.618, 110291.276
attribute	ACCELERATION_T4ms	bcodmo_name	String	unknown
attribute	ACCELERATION_T4ms	description	String	acceleration of fish at t4ms
attribute	ACCELERATION_T4ms	long_name	String	ACCELERATION T4ms
attribute	ACCELERATION_T4ms	units	String	millimeters/second^2
variable	ACCELERATION_T6ms		double
attribute	ACCELERATION_T6ms	_FillValue	double	NaN
attribute	ACCELERATION_T6ms	actual_range	double	-58536.669, 73398.473
attribute	ACCELERATION_T6ms	bcodmo_name	String	unknown
attribute	ACCELERATION_T6ms	description	String	acceleration of fish at t6ms
attribute	ACCELERATION_T6ms	long_name	String	ACCELERATION T6ms
attribute	ACCELERATION_T6ms	units	String	millimeters/second^2
variable	ACCELERATION_T8ms		double
attribute	ACCELERATION_T8ms	_FillValue	double	NaN
attribute	ACCELERATION_T8ms	actual_range	double	-142550.512, 100571.547
attribute	ACCELERATION_T8ms	bcodmo_name	String	unknown
attribute	ACCELERATION_T8ms	description	String	acceleration of fish at t8ms
attribute	ACCELERATION_T8ms	long_name	String	ACCELERATION T8ms
attribute	ACCELERATION_T8ms	units	String	millimeters/second^2
variable	ACCELERATION_T10ms		double
attribute	ACCELERATION_T10ms	_FillValue	double	NaN
attribute	ACCELERATION_T10ms	actual_range	double	-113445.536, 50527.07
attribute	ACCELERATION_T10ms	bcodmo_name	String	unknown
attribute	ACCELERATION_T10ms	description	String	acceleration of fish at t10ms
attribute	ACCELERATION_T10ms	long_name	String	ACCELERATION T10ms
attribute	ACCELERATION_T10ms	units	String	millimeters/second^2
variable	ACCELERATION_T1point5ms		double
attribute	ACCELERATION_T1point5ms	_FillValue	double	NaN
attribute	ACCELERATION_T1point5ms	actual_range	double	-33419.847, 12211.371
attribute	ACCELERATION_T1point5ms	bcodmo_name	String	unknown
attribute	ACCELERATION_T1point5ms	description	String	acceleration of fish at t12ms
attribute	ACCELERATION_T1point5ms	long_name	String	ACCELERATION T1point5ms
attribute	ACCELERATION_T1point5ms	units	String	millimeters/second^2
variable	ACCELERATION_T16ms		double
attribute	ACCELERATION_T16ms	_FillValue	double	NaN
attribute	ACCELERATION_T16ms	actual_range	double	-15273.842, 10407.529
attribute	ACCELERATION_T16ms	bcodmo_name	String	unknown
attribute	ACCELERATION_T16ms	description	String	acceleration of fish at t14ms
attribute	ACCELERATION_T16ms	long_name	String	ACCELERATION T16ms
attribute	ACCELERATION_T16ms	units	String	millimeters/second^2
variable	ACCELERATION_T20ms		double
attribute	ACCELERATION_T20ms	_FillValue	double	NaN
attribute	ACCELERATION_T20ms	actual_range	double	-12761.829, 9654.676
attribute	ACCELERATION_T20ms	bcodmo_name	String	unknown
attribute	ACCELERATION_T20ms	description	String	acceleration of fish at t16ms
attribute	ACCELERATION_T20ms	long_name	String	ACCELERATION T20ms
attribute	ACCELERATION_T20ms	units	String	millimeters/second^2
variable	CVF_T0		float
attribute	CVF_T0	_FillValue	float	NaN
attribute	CVF_T0	actual_range	float	0.386, 0.821
attribute	CVF_T0	bcodmo_name	String	unknown
attribute	CVF_T0	description	String	chord length-to-fish length ratio at t0
attribute	CVF_T0	long_name	String	CVF T0
attribute	CVF_T0	units	String	dimentionless
variable	CVF_T2ms		float
attribute	CVF_T2ms	_FillValue	float	NaN
attribute	CVF_T2ms	actual_range	float	0.409, 0.842
attribute	CVF_T2ms	bcodmo_name	String	unknown
attribute	CVF_T2ms	description	String	chord length-to-fish length ratio at t2ms
attribute	CVF_T2ms	long_name	String	CVF T2MS
attribute	CVF_T2ms	units	String	dimentionless
variable	CVF_T4ms		float
attribute	CVF_T4ms	_FillValue	float	NaN
attribute	CVF_T4ms	actual_range	float	0.523, 0.881
attribute	CVF_T4ms	bcodmo_name	String	unknown
attribute	CVF_T4ms	description	String	chord length-to-fish length ratio at t4ms
attribute	CVF_T4ms	long_name	String	CVF T4MS
attribute	CVF_T4ms	units	String	dimentionless
variable	CVF_T6ms		float
attribute	CVF_T6ms	_FillValue	float	NaN
attribute	CVF_T6ms	actual_range	float	0.659, 0.978
attribute	CVF_T6ms	bcodmo_name	String	unknown
attribute	CVF_T6ms	description	String	chord length-to-fish length ratio at t6ms
attribute	CVF_T6ms	long_name	String	CVF T6MS
attribute	CVF_T6ms	units	String	dimentionless
variable	CVF_T8ms		float
attribute	CVF_T8ms	_FillValue	float	NaN
attribute	CVF_T8ms	actual_range	float	0.835, 1.004
attribute	CVF_T8ms	bcodmo_name	String	unknown
attribute	CVF_T8ms	description	String	chord length-to-fish length ratio at t8ms
attribute	CVF_T8ms	long_name	String	CVF T8MS
attribute	CVF_T8ms	units	String	dimentionless
variable	CVF_T10ms		float
attribute	CVF_T10ms	_FillValue	float	NaN
attribute	CVF_T10ms	actual_range	float	0.819, 1.006
attribute	CVF_T10ms	bcodmo_name	String	unknown
attribute	CVF_T10ms	description	String	chord length-to-fish length ratio at t10ms
attribute	CVF_T10ms	long_name	String	CVF T10MS
attribute	CVF_T10ms	units	String	dimentionless
variable	CVF_T12ms		float
attribute	CVF_T12ms	_FillValue	float	NaN
attribute	CVF_T12ms	actual_range	float	0.821, 1.0
attribute	CVF_T12ms	bcodmo_name	String	unknown
attribute	CVF_T12ms	description	String	chord length-to-fish length ratio at t12ms
attribute	CVF_T12ms	long_name	String	CVF T12MS
attribute	CVF_T12ms	units	String	dimentionless
variable	CVF_T14ms		float
attribute	CVF_T14ms	_FillValue	float	NaN
attribute	CVF_T14ms	actual_range	float	0.753, 1.014
attribute	CVF_T14ms	bcodmo_name	String	unknown
attribute	CVF_T14ms	description	String	chord length-to-fish length ratio at t14ms
attribute	CVF_T14ms	long_name	String	CVF T14MS
attribute	CVF_T14ms	units	String	dimentionless
variable	CVF_T16ms		float
attribute	CVF_T16ms	_FillValue	float	NaN
attribute	CVF_T16ms	actual_range	float	0.731, 1.027
attribute	CVF_T16ms	bcodmo_name	String	unknown
attribute	CVF_T16ms	description	String	chord length-to-fish length ratio at t16ms
attribute	CVF_T16ms	long_name	String	CVF T16MS
attribute	CVF_T16ms	units	String	dimentionless
variable	CVF_T18ms		float
attribute	CVF_T18ms	_FillValue	float	NaN
attribute	CVF_T18ms	actual_range	float	0.738, 1.001
attribute	CVF_T18ms	bcodmo_name	String	unknown
attribute	CVF_T18ms	description	String	chord length-to-fish length ratio at t18ms
attribute	CVF_T18ms	long_name	String	CVF T18MS
attribute	CVF_T18ms	units	String	dimentionless
variable	CVF_T20ms		float
attribute	CVF_T20ms	_FillValue	float	NaN
attribute	CVF_T20ms	actual_range	float	0.71, 1.02
attribute	CVF_T20ms	bcodmo_name	String	unknown
attribute	CVF_T20ms	description	String	chord length-to-fish length ratio at t20ms
attribute	CVF_T20ms	long_name	String	CVF T20MS
attribute	CVF_T20ms	units	String	dimentionless
variable	INVERSE_RADIUS_T0		float
attribute	INVERSE_RADIUS_T0	_FillValue	float	NaN
attribute	INVERSE_RADIUS_T0	actual_range	float	0.751, 14.71
attribute	INVERSE_RADIUS_T0	bcodmo_name	String	unknown
attribute	INVERSE_RADIUS_T0	description	String	reciprocal radius of inscribed circle in fish's tail-bend; r-1 = v(p / A) at t0
attribute	INVERSE_RADIUS_T0	long_name	String	INVERSE RADIUS T0
attribute	INVERSE_RADIUS_T0	units	String	per millimeter
variable	INVERSE_RADIUS_T2ms		float
attribute	INVERSE_RADIUS_T2ms	_FillValue	float	NaN
attribute	INVERSE_RADIUS_T2ms	actual_range	float	0.75, 9.008
attribute	INVERSE_RADIUS_T2ms	bcodmo_name	String	unknown
attribute	INVERSE_RADIUS_T2ms	description	String	reciprocal radius of inscribed circle in fish's tail-bend; r-1 = v(p / A) at t2ms
attribute	INVERSE_RADIUS_T2ms	long_name	String	INVERSE RADIUS T2ms
attribute	INVERSE_RADIUS_T2ms	units	String	per millimeter
variable	INVERSE_RADIUS_T4ms		float
attribute	INVERSE_RADIUS_T4ms	_FillValue	float	NaN
attribute	INVERSE_RADIUS_T4ms	actual_range	float	0.696, 5.174
attribute	INVERSE_RADIUS_T4ms	bcodmo_name	String	unknown
attribute	INVERSE_RADIUS_T4ms	description	String	reciprocal radius of inscribed circle in fish's tail-bend; r-1 = v(p / A) at t4ms
attribute	INVERSE_RADIUS_T4ms	long_name	String	INVERSE RADIUS T4ms
attribute	INVERSE_RADIUS_T4ms	units	String	per millimeter
variable	INVERSE_RADIUS_T6ms		float
attribute	INVERSE_RADIUS_T6ms	_FillValue	float	NaN
attribute	INVERSE_RADIUS_T6ms	actual_range	float	0.476, 2.996
attribute	INVERSE_RADIUS_T6ms	bcodmo_name	String	unknown
attribute	INVERSE_RADIUS_T6ms	description	String	reciprocal radius of inscribed circle in fish's tail-bend; r-1 = v(p / A) at t6ms
attribute	INVERSE_RADIUS_T6ms	long_name	String	INVERSE RADIUS T6ms
attribute	INVERSE_RADIUS_T6ms	units	String	per millimeter
variable	INVERSE_RADIUS_T8ms		float
attribute	INVERSE_RADIUS_T8ms	_FillValue	float	NaN
attribute	INVERSE_RADIUS_T8ms	actual_range	float	0.208, 2.5
attribute	INVERSE_RADIUS_T8ms	bcodmo_name	String	unknown
attribute	INVERSE_RADIUS_T8ms	description	String	reciprocal radius of inscribed circle in fish's tail-bend; r-1 = v(p / A) at t8ms
attribute	INVERSE_RADIUS_T8ms	long_name	String	INVERSE RADIUS T8ms
attribute	INVERSE_RADIUS_T8ms	units	String	per millimeter

The information in the table above is also available in other file formats (.csv, .htmlTable, .itx, .json, .jsonlCSV1, .jsonlCSV, .jsonlKVP, .mat, .nc, .nccsv, .tsv, .xhtml) via a RESTful web service.